2021 WORLDSymposium Ayuko Iverson, Narita Aya, Handover to Otake, GBA-PD penetrance, Mutation GBA-PD

2021 WORLDSymposium Ayuko Iverson

GD genotype and Tx status had no effect on development or progression of PD. AAO is earlier (54.6 vs 60 (historical)) but variable (41-83 range)

Narita Aya (Iottori / Tottori univ)

itemnote / readoutadditional note
GDPatient stratificationPatients with Epilepsy
ResponsiveSymptom (sz?), age?
GBA activity improvement - phenotype improvement relationshippaper에는 무관으로 보임.venglustat에선, 6개월이나 GBA activity 증가시켰는데, saccade 불변, 왜?
PD markerEEG
Oculomotor signsHow long
NHP?
In vitro screeningfibroblastPassage number
Total time (1-2 weeks?)
Will reflect CNS?
PBMCWill reflect CNS?

Handover to Otake on GD

Candidate BMsRelated activitiesDocuments
Brain volumeSanofi's venglustat measured this in P2 program using vMRI2020 & 2021 poster
Brain regional connectivitySanofi's venglustat measured this in P2 program using fMRI2020 & 2021 poster
Saccadic eye movementSanofi's venglustat measured this in P2 program2019 poster
PLR and other retinal signs Dr. Narita (Tottori univ) is proposing PLR as an important novel biomarker (2016 Narita's paper). We had a meeting with her thanks to Hirozane san's arrangement.
Tomimatsu san's group have already budgeted for purchasing animal PLR detector for FY2021.
In nGD there are other pathological retinal changes (eg. retinal thinning) and these may be measured by OCT both in mice and human.
- We were going to meet with an ophthalmologist (Dr. Ishikawa of Kitasato University) for consultation as per Dr. Narita's recommendation. But I didn't proceed with this as GBA activator was terminated.
- RDTM was going to measure retinal histology in GD mice but didn't proceed because, unlike ND DDU's D409V mice, their D409V mice didn't show robust GD phenotype (eg. GlcCer and GlcSph).
Meeting summary (20200901)

GBA-PD / Penetrance (LIFETIME)

citationGBA hetero penetranceGBA hetero contextLifetime Relative Risk (RR)GBA homo penetrance
(McNeill et al. 2012, PMID 22577228)15%Family Hx of PD30
(Alcalay et al. 2014, PMID 24756352)7.7%Regardless of Family Hx of PD9.1%
(Anheim et al. 2012, PMID 22282650)29.7%Family Hx of PD

Mutation GBA-PD

(Parlar, 2023) total 371 variants ( GBA-PD Browser 20240313 406) https://pdgenetics.shinyapps.io/gba1browser/

Classification

severemildriskUnknown
# of variants84223262
% of 37122.6%5.9%0.8%70.6%
GDCause Type ICause Type II OR IIIXimpossible to determine if they are severe or mild based on GD (causing GD임은 앎, though)
PD↑ as a group↑ as a group↑ PDX (not statistically associated with PD)
representative variantsp.L444Pp.N370Sp.E326K and p.T369Mp.R44C: in PD there are contradicting evidence for it being a risk factor, as one study reported an odds ratio of 18 in one study with very large confidence intervals,40 while in another study it was found more frequently in controls.41
OR for PD>10>2<2

Three most common variants in decreasing order

  • (2019 Blauwendraat): (p.E326K, p.T369M and p.N370S)
  • (Huh, 2020 #1105): p.E326K, p.N370S, p.T369M
  • PPMI:? 내 DATSCAN 분석: (p.E326K, p.T369M and p.N370S)
  • (Davis, 2016 #1247) p.E326K, p.N370S, L444P, p.T369M

Severe = cause GD type 3 = neuronopathic

  • P.L444P: p.Leu483Pro p.L483P (rs421016, UKBBN에 0개) c.1448T>C (이것 뭐?) (Alfonso, 2005 #599) 22-fold reduction in the activity of the N370S enzyme, L444P has a lower stability than the wild-type protein [13].
  • 84GG (The insertion of a single nucleotide, a second guanine at cDNA nt 84, null mutation임): c.84dupG
  • D409H
  • IVS1011G>T: splicing mutation
  • G195E, H255Q, R257Q, P266L, R359X, G377S, , L444R, A456P, N462K, R120W, and R463C, p.G377S,

Mild = cause GD type I = non-neuronopathic

LocirsNoAlleleFrequencyEffect on GRSKnown/nearest genes
1:155235843rs76763715 UKBBN 에 4개보이나 모두 정상 PN3705T>C, G0.0071.22GBA:N370S

P.N370S: p.N409S (=rs76763715), c.1226A>G (Alfonso, 2005 #599) p.Asn409Ser, asparagine (이러한 asparagine 같은 것들이 three aa 일걸, 그래서 이런 mutation 이 three sequence change 일걸) residue at position 370 is thought to have an indirect role in maintaining an active conformation, Ohashi et al. demonstrated that the N370S mutation has an abnormal catalytic site, because it differs from the normal enzyme in its inhibition by both conduritol B epoxide and glucosphingosine

IVS2 G+1----A+1: c.115+1G>A

T408S, ((He and Grabowski 1992, PMID 1415223)

Other risk variant (can’t be classified into mild or severe because these don’t cause GD)

p.E326K (p.Glu365Lys) → ↑ PD (2018 Leonard, 2018 (Huang, 2018 #738) Huang with OR of 1.99),

(2018 Huang: this is the same as p.E365K,)

(2016 Davis) E326K is a nonconserved residue and is not predicted to significantly alter glucocerebrosidase enzyme activity in silico, several studies expressing GBA constructs with E326K suggest that this polymorphism reduces enzyme activity. 27,30,31

↓ gba activity in hetero-PD Brain (Nelson et al. 2018, PMID 29196214), 여기 원문에서 잘 확인이 안되는데?,

↓ gba activity in dried blood spots (E326K, P = 0.009; T369M, P50.001).(Alcalay, 2015 #258, table3)

LocirsNoAlleleFrequencyEffect on GRSKnown/nearest genes
1:155236376rs2230288 UKBBN 에 12 개중 hetero 하나발견G>A, C>T0.0260.74GBA:E365K = E326K

p.T369M (p.Thr408Met, =p.T408M, = rs75548401) G>A, UKBBN에 6개중

The minor allele of GBA p.T408M (p = 4.9 x is (certainly) associated with increased PD risk).(Benitez et al. 2016, PMID)

GBA Activity: (Alcalay, 2015 #258) ↓ in patients with PD and controls combined (n=9), compared with that in noncarriers (table 3, 7.64 vs 11.93 mmol/L/h, p, 0.001), in dried blood spot

(Iwaki et al. 2019, PMID)

R39C 도 이런 것 같은데?

Unknown function

p.E388K Excluded in (Cilia et al. 2016, PMID 27632223)

Four major GBA variants in GBA-PD

variantFrequency in GBA-PD (Malek, 2018 #549)Cause GD?Increase PD risk?Reduce GBA activity?
L444P15.5%YesYesYes
N370S5.7%YesYesYes
E326K (=p.E365K)44.6%NoYes (Huang, 2018 #738) (OR of 1.99),Yes (Alcalay, 2015 #258)
T369M (=p.T408M),18.1%NoYes (Benitez et al., 2016)Yes (Alcalay, 2015 #258)

Variant position table (hg38)

Protein codersIDPos in hg38
L444Prs421016chr1:155235252
H255Qrs367968666chr1:155237458
W291XNANA
R131Lrs398123530chr1:155238596
D409Hrs1064651chr1:155235727
N370Srs76763715chr1:155235843
E326Krs2230288chr1:155236376
T369Mrs75548401chr1:155236246
W378GNANA
R2INANA
R501C
N227Srs535896234chr1:155238215
Y212Hrs121908300chr1:155238144

Postmortem of GBA-PD

Good postmortem review on (Schneider, 2017 #1056): key finding is ↑ cortical LBs

citationGBALBDnote
(... et al. 2004, ...15234332)HomozPDDGD+PDD (n=4), GD only etc
[GD+PDD Patients (n=4)] Diffuse and not limited in brain stem and limbic system (ie LB pathology in hippocampus and cortex with astrogliosis)
(...ka et al. 2011, ...20971030)!this is not a post-mortem
(...i et al. 2003, ...12809640)HomozPDGD+PD

Uncertain Spans

locationtranscriptionuncertainty
E326K noteP50.001reads as written; OCR may have misread P=0.001 as P50.001.
Postmortem tableleftmost citation column truncatedthe citation prefixes are clipped; preserved as ... placeholders.